[1] Blakemore R. Magnetotactic bacteria[J]. Science, 1975, 190(4212): 377-379. [2] Pan Y X,Lin W,Li J H, et al. Reduced efficiency of magnetotaxis in magnetotactic coccoid bacteria in higher than geomagnetic fields[J]. Biophys J, 2009, 97(4): 986-991. [3] Thornhill RH,Grant Burgess J,Sakaguchi T, et al. A morphological classification of bacteria containing bullet-shaped magnetic particles[J]. FEMS Microbiology Letters, 1994, 115(2-3): 169-176. [4] Vali H,F?rster O,Amarantidis G, et al. Magnetotactic bacteria and their magnetofossils in sediments[J]. Earth and Planetary Science Letters, 1987, 86(2-4): 389-400. [5] Flies C,Peplies J,Schüler D. Combined approach for characterization of uncultivated magnetotactic bacteria from various aquatic environments[J]. Applied and Environmental Microbiology, 2005, 71(5): 2723-2731. [6] Lin W,Pan Y X. Uncultivated magnetotactic cocci from yuandadu park in beijing, china[J]. Appl Environ Microbiol, 2009, 75(12): 4046-4052. [7] Faivre D,Schüler D. Magnetotactic bacteria and magnetosomes[J]. Chem Rev, 2008, 108(11): 4875-4898. [8] Frankel RB,Blakemore R. Navigational compass in magnetic bacteria[J]. Journal of Magnetism and Magnetic Materials, 1980, 15: 1562-1564. [9] Winklhofer M,Petersen N. (Paleomagnetism and magnetic bacteria. In:Schüler D. ed). Magnetoreception and magnetosomes in bacteria[M],Heidelberg: Springer,2007:255-273. [10] Scheffel A,Gruska M,Faivre D, et al. An acidic protein aligns magnetosomes along a filamentous structure in magnetotactic bacteria[J]. Nature, 2006, 440(7080): 110-114. [11] Komeili A,Li Z,Newman DK, et al. Magnetosomes are cell membrane invaginations organized by the actin-like protein MamK[J]. Science, 2006, 311(5758): 242-245. [12] Komeili A,Vali H,Beveridge TJ, et al. Magnetosome vesicles are present before magnetite formation, and MamA is required for their activation[J]. Proceedings of the National Academy of Sciences, 2004, 101(11): 3839-3844. [13] Yamamoto D,Taoka A,Uchihashi T, et al. Visualization and structural analysis of the bacterial magnetic organelle magnetosome using atomic force microscopy[J]. Proceedings of the National Academy of Sciences, 2010, 107(20): 9382-9387. [14] Murat D,Quinlan A,Vali H, et al. Comprehensive genetic dissection of the magnetosome gene island reveals the step-wise assembly of a prokaryotic organelle[J]. Proceedings of the National Academy of Sciences, 2010, 107(12): 5593-5598. [15] Frankel RB,Bazylinski DA. How magnetotactic bacteria make magnetosomes queue up[J]. Trends Microbiol, 2006, 14(8): 329-331. [16] Matsunaga T,Okamura Y,Fukuda Y, et al. Complete genome sequence of the facultative anaerobic magnetotactic bacterium Magnetospirillum sp. strain AMB-1[J]. DNA Res, 2005, 12(3): 157-166. [17] Schübbe S,Wurdemann C,Peplies J, et al. Transcriptional organization and regulation of magnetosome operons in Magnetospirillum gryphiswaldense[J]. Appl Environ Microbiol, 2006, 72(9): 5757-5765. [18] Grünberg K,Müller EC,Otto A, et al. Biochemical and Proteomic Analysis of the Magnetosome Membrane in Magnetospirillum gryphiswaldense[J]. Applied and Environmental Microbiology, 2004, 70(2): 1040-1050. [19] Scheffel A,Schüler D. The Acidic Repetitive Domain of the Magnetospirillum gryphiswaldense MamJ Protein Displays Hypervariability but Is Not Required for Magnetosome Chain Assembly[J]. Journal of Bacteriology, 2007, 189(17): 6437-6446. [20] Bork P,Sander C,Valencia A. An ATPase domain common to prokaryotic cell cycle proteins, sugar kinases, actin, and hsp70 heat shock proteins[J]. Proceedings of the National Academy of Sciences, 1992, 89(16): 7290-7294. [21] Gitai Z. Diversification and specialization of the bacterial cytoskeleton[J]. Current opinion in cell biology, 2007, 19(1): 5-12. [22] Taoka A,Asada R,Wu LF, et al. Polymerization of the Actin-Like Protein MamK, Which Is Associated with Magnetosomes[J]. Journal of Bacteriology, 2007, 189(23): 8737-8740. [23] Pradel N,Santini CL,Bernadac A, et al. Biogenesis of actin-like bacterial cytoskeletal filaments destined for positioning prokaryotic magnetic organelles[J]. Proceedings of the National Academy of Sciences, 2006, 103(46): 17485-17489. [24] Olga Draper,Meghan Byrne,Sepehr Keyhani, et al., In vivo bacterial actin filament dynamics in Magnetospirillum magneticum sp. AMB-1 In International Symposium on Magnetotactic Bacteria and Biomineralization[C], Pan Y X, et al. Beijing, (2010); p 25. [25] Rioux JB,Philippe N,Pereira S, et al. A second actin-like MamK protein in Magnetospirillum magneticum AMB-1 encoded outside the genomic magnetosome island[J]. PLoS ONE, 2010, 5(2): e9151. [26] Taoka A,Asada R,Sasaki H, et al. Spatial localizations of Mam22 and Mam12 in the magnetosomes of Magnetospirillum magnetotacticum[J]. J Bacteriol, 2006, 188(11): 3805-3812. [27] Zeytuni N,Zarivach R. Crystallization and preliminary crystallographic analysis of the Magnetospirillum magneticum AMB-1 and M. gryphiswaldense MSR-1 magnetosome-associated proteins MamA[J]. Acta Crystallographica Section F: Structural Biology and Crystallization Communications, 2010, 66(7): 824-827. [28] Quinlan A,Murat D,Vali H, et al. The HtrA/DegP family protease MamE is a bifunctional protein with roles in magnetosome protein localization and magnetite biomineralization[J]. Mol Microbiol, 2011. [29] Tanaka M,Arakaki A,Matsunaga T. Identification and functional characterization of liposome tubulation protein from magnetotactic bacteria[J]. Mol Microbiol, 2010, 76(2): 480-488. [30] Li J H,Pan Y X,Chen G J, et al. Magnetite magnetosome and fragmental chain formation of Magnetospirillum magneticum AMB-1: transmission electron microscopy and magnetic observations[J]. Geophysical Journal International, 2009, 177(1): 33-42. [31] Schübbe S,Kube M,Scheffel A, et al. Characterization of a Spontaneous Nonmagnetic Mutant of Magnetospirillum gryphiswaldense Reveals a Large Deletion Comprising a Putative Magnetosome Island[J]. Journal of Bacteriology, 2003, 185(19): 5779-5790. [32] Katzmann E,Scheffel A,Gruska M, et al. Loss of the actin-like protein MamK has pleiotropic effects on magnetosome formation and chain assembly in Magnetospirillum gryphiswaldense[J]. Mol Microbiol, 2010, 77(1): 208-224. [33] F. D. Müller,M. Messerer,S. Ullrich, et al., Functional analysis of cytoskeletal proteins implicated in magnetosome formation in M.gryphiswaldense, In International Symposium on Magnetotactic Bacteria and Biomineralization[C], Pan Y X, et al. Beijing, (2010); pp 45-46. [34] Graumann P. Cytoskeletal elements in bacteria[J]. Current opinion in microbiology, 2004, 7(6): 565-571. [35] Bi E,Lutkenhaus J. FtsZ ring structure associated with division in Escherichia coli[J]. Nature, 1991, 354: 161-164. [36] Ding Y,Li J L,Liu J N, et al. Deletion of the ftsZ-like gene results in the production of superparamagnetic magnetite magnetosomes in Magnetospirillum gryphiswaldense[J]. J Bacteriol, 2010, 192(4): 1097-1105. [37] Tanaka M,Okamura Y,Arakaki A, et al. Origin of magnetosome membrane: Proteomic analysis of magnetosome membrane and comparison with cytoplasmic membrane[J]. Proteomics, 2006, 6(19): 5234-5247. [38] Richter M,Kube M,Bazylinski DA, et al. Comparative genome analysis of four magnetotactic bacteria reveals a complex set of group-specific genes implicated in magnetosome biomineralization and function[J]. J Bacteriol, 2007, 189(13): 4899-4910. [39] Schübbe S,Williams T,Xie G, et al. Complete genome sequence of the chemolithoautotrophic marine magnetotactic coccus strain MC-1[J]. Applied and Environmental Microbiology, 2009, 75(14): 4835-4852. [40] Nakazawa H,Arakaki A,Narita-Yamada S, et al. Whole genome sequence of Desulfovibrio magneticus strain RS-1 revealed common gene clusters in magnetotactic bacteria[J]. Genome Res, 2009, 19(10): 1801-1808. [41] Jogler C,Wanner G,Kolinko S, et al. Conservation of proteobacterial magnetosome genes and structures in an uncultivated member of the deep-branching Nitrospira phylum[J]. Proceedings of the National Academy of Sciences, 2010, 180(3):1134-1139. [42] Jogler C,Kube M,Schübbe S, et al. Comparative analysis of magnetosome gene clusters in magnetotactic bacteria provides further evidence for horizontal gene transfer[J]. Environmental Microbiology, 2009, 11(5): 1267-1277.